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TERRA: Terrain-Aware Reconstruction, Retargeting and Control for Musculoskeletal Locomotion
Merkourios Simos, Chengkun Li, Bianca Ziliotto, Alexander Mathis
Intelligence
Status: succeeded | Model: Gemma-4-26B-A4B | Prompt: intel-v1 | Confidence: 89%
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Summary
The paper introduces TERRA, an end-to-end pipeline for terrain-aware reconstruction, retargeting, and control of musculoskeletal locomotion. It addresses the challenge of training muscle-actuated agents on non-flat terrain by reconstructing support geometry from kinematic data alone using terrain priors and contact evidence. TERRA then retargets human motion to a complex musculoskeletal model (MyoFullBody) while respecting anatomical and contact constraints. The resulting motion-terrain pairs are used to train a single reinforcement learning policy capable of diverse locomotion tasks across ramps, stairs, platforms, and seats.
Entities (14)
Relation Signals (8)
TERRA â retargets â MyoFullBody
confidence 95% · TERRA further considers anatomical, tendon-continuity, and contact constraints during retargeting... to the musculoskeletal model.
TERRA â trains â muscle-actuated control policy
confidence 95% · Using the resulting motion-terrain pairs from five datasets, we successfully train a single muscle-actuated control policy
TERRA â uses â terrain priors
confidence 95% · TERRA combines terrain priors, estimated contacts, and negative free-space evidence to recover task-relevant support geometry.
TERRA â adapts â OmniRetarget
confidence 90% · TERRA adapts OmniRetargetâs interaction-mesh representation and adds target-specific anatomical and terrain-interaction terms
TERRA â improves â terrain accuracy
confidence 90% · TERRA improves terrain accuracy, sharply reduces anatomical and interaction violations
MyoFullBody â isconfiguredfrom â MuscleMimic
confidence 90% · We used the finger-disabled MyoFullBody configuration from MuscleMimic [7]
TERRA â supports â AMASS
confidence 90% · TERRA accepts motion inputs in AMASS-compatible SMPL-H format
TERRA â outperforms â SceneBot
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Abstract
Abstract:Recent advances in musculoskeletal modeling and reinforcement learning have enabled muscle-actuated agents to reproduce increasingly complex human motions. Yet these capabilities remain largely confined to flat ground, in part because motion datasets rarely include aligned terrain geometry and because retargeting terrain interactions to complex musculoskeletal bodies is challenging. We present TERRA, an end-to-end pipeline for terrain-aware retargeting and control of musculoskeletal locomotion. From kinematic trajectories alone, TERRA combines terrain priors, estimated contacts, and negative free-space evidence to recover task-relevant support geometry. TERRA further considers anatomical, tendon-continuity, and contact constraints during retargeting. Using the resulting motion-terrain pairs from five datasets, we successfully train a single muscle-actuated control policy on 9.4 hours of diverse locomotion. Across reconstruction, retargeting, and held-out tracking benchmarks, TERRA improves terrain accuracy, sharply reduces anatomical and interaction violations, and achieves the highest observed completion rate over supported terrain families. Overall, TERRA provides a practical route from scene-less motion data to muscle-actuated locomotion over diverse non-flat terrain. Project website: this https URL
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- Source: https://arxiv.org/abs/2609.38653v1
- Canonical: https://arxiv.org/abs/2609.38653v1
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TERRA: Terrain-Aware Reconstruction, Retargeting and Control for Musculoskeletal Locomotion Merkourios Simos Affiliation: EPFL Chengkun Li Affiliation: EPFL Bianca Ziliotto Affiliation: EPFL Alexander Mathis â thanks: Contact: alexander.mathis@epfl.châ thanks: This work was supported by Swiss National Science Foundation (SNSF) (310030_212516), the Simons foundation (SFI-AN-NC-SCN-00007276-14), and a Boehringer Ingelheim Fonds PhD stipend (B.Z.). M.S. acknowledges the support of the Onassis Foundation Scholarshipsâ Program. Affiliation: EPFL Abstract Recent advances in musculoskeletal modeling and reinforcement learning have enabled muscle-actuated agents to reproduce increasingly complex human motions. Yet these capabilities remain largely confined to flat ground, in part because motion datasets rarely include aligned terrain geometry and because retargeting terrain interactions to complex musculoskeletal bodies is challenging. We present TERRA, an end-to-end pipeline for terrain-aware retargeting and control of musculoskeletal locomotion. From kinematic trajectories alone, TERRA combines terrain priors, estimated contacts, and negative free-space evidence to recover task-relevant support geometry. TERRA further considers anatomical, tendon-continuity, and contact constraints during retargeting. Using the resulting motion-terrain pairs from five datasets, we successfully train a single muscle-actuated control policy on 9.4 hours of diverse locomotion. Across reconstruction, retargeting, and held-out tracking benchmarks, TERRA improves terrain accuracy, sharply reduces anatomical and interaction violations, and achieves the highest observed completion rate over supported terrain families. Overall, TERRA provides a practical route from scene-less motion data to muscle-actuated locomotion over diverse non-flat terrain. Project website: https://cnai.epfl.ch/terra/ â aftertitle: Fig. 1: Representative outputs from TERRA. Each tile shows a motion-derived static collision terrain and the corresponding MyoFullBody reference used for muscle-driven tracking. The supported terrain families (affordances) are ramps, staircases, independent horizontal supports, and seats. I Introduction Whether trail-running on Mont Blanc, climbing a flight of stairs, or simply sitting down in a chair, humans need to coordinate hundreds of muscles as the environment and the underlying terrain change. Understanding how such control arises across diverse affordances [1] is a fundamental problem in neuroscience [2] and an increasingly practical one in robotics, as humanoids must navigate the same spaces designed for humans. Physics-based musculoskeletal models combined with reinforcement learning (RL) provide a powerful framework for studying how complex movement can emerge from muscle-level control. Early work used task-driven objectives to generate individual skills such as walking and running [3, 4]. More recent approaches leveraged large-scale motion-capture datasets and motion imitation to learn broad behavioral repertoires that can be reused for downstream tasks [5, 6, 7, 8, 9]. However, the behavioral repertoires remain almost entirely confined to flat ground. Two challenges make non-flat locomotion particularly difficult. First, existing motion-capture datasets rarely provide aligned terrain geometry. Second, transferring human motion to a complex musculoskeletal body requires more than matching joint positions: the retargeted motion must preserve contacts, avoid penetration, foot skating, and floating, and remain compatible with anatomical joint and musculotendon limits. These errors are amplified by non-flat terrain interaction and can render a reference motion infeasible (Table I). We present TERRA, an end-to-end framework for terrain-aware musculoskeletal retargeting and control. From kinematics alone, TERRA reconstructs motion-relevant support geometry by combining terrain priors with estimated contact events, foot orientation, and free-space evidence from the moving body. It then extends interaction-mesh retargeting with anatomical, contact, clearance, and collision constraints to produce valid references for a full-body, muscle-actuated model. The resulting motionâterrain pairs form a 9.4-hour, five-source library for a single reference-conditioned controller [10, 11, 12, 13, 14]. Across ramps, stairs, platforms, and seats, TERRA reconstructs motion-supported terrain, reduces reference violations, and enables one controller to complete held-out motions from all four families. Lastly, we qualitatively compare generated muscle activity with held-out EMG and vertical ground-reaction-forces (GRF) across flat ground, ramps, stairs, and sit/stand motions, characterizing similarities and differences in normalized waveform shape. Code and data will be made publicly available. I Related Work Motion-conditioned terrain reconstruction. Several recent methods recover or synthesize surrounding scene structure based on motion trajectories, using learned contact and free-space cues or physics-based interaction constraints [15, 16]. Complementary video-based pipelines jointly reconstruct human motion and scene geometry but rely on visual observations [17, 18, 19]. Without visual scene observations, TIP jointly estimates inertial motion and a local terrain height field [20]. Most closely related, SceneBot constructs contact-rich environments from retargeted motion by placing and merging constant-height terrain patches [21]. TERRA instead performs structured inference over explicit terrain priors, including platforms, stair flights, inclined ramps, and seated supports. Whereas SceneBot evaluates its reconstructed scenes primarily through downstream tracking success, TERRA additionally measures geometry against paired ground-truth terrain. Because SceneBotâs code is not publicly available, we compare against an explicit constant-height patch baseline adapted from its method and TIP. Motion retargeting. Large-scale human-motion repositories have made motion imitation a viable and scalable strategy for training general-purpose humanoid controllers [10, 22, 23]. Recorded motions must first be mapped into a robotâs morphology. Existing methods include keypoint-based optimization supporting diverse embodiments [24], and learned cross-morphology mappings [25]. Recent work has also incorporated physical feasibility [23, 26]. Lastly, OmniRetarget proposed a shared body-scene interaction mesh that preserves interactions with objects and terrain [27]. While OmniRetarget assumes access to the terrain or object geometry used to construct that interaction mesh, TERRA addresses a complementary upstream problem: it recovers task-relevant support geometry when a motion dataset provides body kinematics without a scene model. TERRA adapts OmniRetargetâs interaction-mesh representation and adds target-specific anatomical and terrain-interaction terms for a muscle-driven embodiment. Musculoskeletal control. Advances in musculoskeletal modeling and simulation have made physiologically detailed, contact-rich bodies increasingly accessible to learning-based control [28, 29, 30, 31]. Several works tackled the challenge of efficient exploration in high-dimensional muscle actuation space [32, 33, 34], enabling multi-task control of increasing dexterity and even athleticism [35, 36, 8, 37]. More recently, motion imitation has enabled reusable locomotor repertoires, empirical validation of simulated muscle activity, and universal policies controlling muscle-actuated bodies across hundreds of reference motions [5, 6, 7, 9]. Nevertheless, musculoskeletal imitation remained largely confined to flat ground. TERRA extends musculoskeletal imitation learning to three-dimensional locomotion by training a single muscle-actuated policy on motions paired with reconstructed terrain. TABLE I: Capabilities of representative motion-processing methods. MSK denotes support for musculoskeletal target models. Method Kinematic constraints Terrain reconstruction Terrain retargeting MSK GMR [24] Joint bounds â â â OmniRetarget [27] Hard interaction â â â SceneBot [21] Joint bounds â â â MuscleMimic [7] Joint/equality â â â TERRA Hard + anatomy â â â Fig. 2: TERRA method overview. Heterogeneous motion data are converted to SMPL-H, used to reconstruct compact collision terrain, and retargeted to the musculoskeletal model with terrain-aware constraints. The resulting paired trajectoryâterrain scenes train a muscle-actuated tracking policy, whose biomechanical outputs are compared with subject-matched force-plate and EMG data on held-out motions. I Method We first introduce the musculoskeletal model, source motion, and terrain priors. We then describe the terrain reconstruction method, the retargeting stage, musculoskeletal controller training, and the biomechanical comparison pipeline (Fig. 2). I-A Problem Setup Musculoskeletal model. We used the finger-disabled MyoFullBody configuration from MuscleMimic [7], with 354 Hill-type musculotendon actuators and 83 MuJoCo joints: one floating-root joint and 82 scalar articulated joints. The model also contains 49 active polynomial couplings for dependent lumbar, scapulohumeral, and knee joints. In order to align the model with reference motion data, we paired K=17K=17 SMPL-H [38] landmarks with MyoFullBody body origins: pelvis, spine, and head; bilateral hip, knee, ankle, and toe; and bilateral shoulder, elbow, and wrist. We denote their target-model positions by iâ() p_i( q). Alignment was performed by placing MyoFullBody and SMPL-H in corresponding neutral poses, optimizing the SMPL-H shape coefficients and global scale, and applying residual position offsets to source motions. Collision geometries in the feet, thighs, shanks, and posterior pelvis were modeled as capsule and ellipsoid primitives. Source motion. TERRA accepts motion inputs in AMASS-compatible SMPL-H format [10]. For marker-based datasets, TERRA additionally supports C3D, MAT, and TRC formats (Fig. 2). Each sequence is transformed into 52 world-space joint positions and rotations using the SMPL-H shape fitted to MyoFullBody, then translated vertically so that the lowest fitted toe lies at z=0z=0. Terrain priors. TERRA defines four terrain priors: independent support boxes, inclined ramps, staircases, and seat supports (Fig. 2). Each prior is implemented as an assembly of finite, static MuJoCo box collision geometries. Independent supports and seats use horizontal boxes, staircases use an ordered set of horizontal boxes, and ramps use a pitched box with a planar top face. All terrain boxes use the same contact and friction parameters as the floor. I-B Terrain Reconstruction TERRA constructs terrain directly from motion trajectories via terrain priors. Intervals in which an ankle or toe landmark remains locally stationary and low provide positive surface evidence, while the remaining body trajectory provides free-space evidence that bounds the admissible geometry. TERRA distinguishes discrete horizontal surfaces from continuous ramps using two additional motion cues: the neutral-calibrated ankleâtoe orientation during stance and the swing-foot clearance profile between consecutive stance intervals of the same foot. Support extraction and calibration. For each left and right ankle and toe landmark j, we define candidate stationary intervals e as contiguous runs satisfying â„Ëj,tâ„2<vs x_j,t _2<v_s for all tâetâ e. A candidate is retained when |e|â„nmin,zÂŻeâ€minÏâeâĄzj,Ï+Ï”loc,|e|â„ n_ , z_e†_Ï _ez_j,Ï+ _loc, (1) where zÂŻe z_e is the median landmark height and eW_e is a local temporal window. Candidate ankle and toe intervals that overlap with a detected seated interval are excluded from surface-height estimation. To account for the distance between the joint centers and the bodyâs contact surface, we subtract a joint-specific offset ojo_j and treat h^e=zÂŻeâojâĄ(e) h_e= z_e-o_j(e) as surface height. When available, offsets come from a subject-matched flat reference; otherwise, we estimate them from self-calibration for datasets with a flat portion, or from the lowest motion-supported level. Distinct surface-height levels are computed by sorting h^e h_e and starting a new level whenever adjacent values differ by more than 44 cm. Any initial group spanning more than 88 cm is recursively divided at its largest internal gap. The height of each level is computed by first averaging h^e h_e within each landmark (e.g. left toe) and then averaging across landmarks. Continuous versus discrete terrain. Let u denote the projection of each stationary intervalâs median horizontal location onto the first principal direction a of these locations, oriented toward increasing height. We fit the flatâinclineâflat profile hâĄ(u)=h0+sâĄ[clipâĄ(u,u0,u1)âu0]h(u)=h_0+s\,[clip(u,u_0,u_1)-u_0] (2) where h0h_0 is the lower landing height, s is the incline grade, and u0u_0 and u1u_1 are the coordinates at which the incline begins and ends. Let Ξt _t be the ankleâtoe pitch, Ξâ0 _ ^0 its neutral value for foot â , and Îłt _t the angle between the foot heading and ramp direction. Using medians first within each stationary interval and then across intervals, we compute Eflat E_flat =medâĄ|ΞtâΞâ0|, =med| _t- _ ^0|, (3) Eramp E_ramp =med|ΞtâΞâ0âarctan(scosÎłt)|. =med| _t- _ ^0- (s _t)|. The smaller error determines whether the foot orientation better matches a flat or inclined surface. When |EflatâEramp|â€2â|E_flat-E_ramp|†2 , a normalized swing-clearance score compares the foot trajectory with a linear height transition and assigns near-linear trajectories to ramps and trajectories with early ascent or delayed descent to discrete surfaces. The 2â2 ambiguity band corresponds to approximately 7-m endpoint uncertainty over a 20-cm foot chord; the swing cue must exceed 5% of the observed support-level change. Discrete terrain geometry. For motions containing at least three raised surface-height levels, a staircase candidate with a common horizontal axis and width with fitted riser height is considered. An independent-box candidate is also constructed. The yaw of each raised level is aligned with the principal axis of its horizontal contact points, and spatially separated contacts on the same level are assigned to different boxes. Each contact envelope is expanded by a foot-sized margin, then trimmed where the resulting geometry would intersect free-space body trajectories. A staircase prior is selected when candidate treads are within 55 cm of every raised stationary foot and no other body landmark penetrates it by more than 55 cm; otherwise, terrain is constructed with independent boxes. Seat reconstruction. Seats are represented by boxes and detected when pelvis speed remains below 0.150.15 m/s for at least 0.40.4 s and its median horizontal position lies at least 0.100.10 m outside the convex hull of the feet and grounded limb landmarks. The seat top is the median of the 1010th-percentile heights of the robot-fitted SMPL-H posterior pelvis/hip surface at the first, middle, and last interval frames. Candidates outside the 0.040.04â0.600.60 m height range are discarded. I-C Terrain-Aware Motion Retargeting Once a terrain is constructed, TERRA retargets the paired motion by adapting OmniRetargetâs interaction-mesh objective, adding target-specific anatomical and terrain-interaction terms, and solving the resulting objective by sequential quadratic programming. Interaction mesh. Following OmniRetarget [27], let i,t x_i,t be the source landmark corresponding to the target-model position iâ() p_i( q), and let S=[1,âŠ,NS]â€S=[ s_1,âŠ, s_N_S] contain samples of the reconstructed scene. We sample each box top with spacing no greater than 0.200.20 m and add a coarse floor grid after removing points inside box footprints. At each frame, the source and robot vertex matrices are V~t V_t =[1,tâŻK,tS], = [ x_1,t T\ ·s\ x_K,t T\ S T ] T, (4) Vtâ() V_t( q) =[1()âŻK()S]. = [ p_1( q) T\ ·s\ p_K( q) T\ S T ] T. A Delaunay tetrahedralization of V~t V_t defines neighbor sets i,tN_i,t and the uniform Laplacian (LtâV)i=iâ|i,t|â1ââjâi,tj(L_tV)_i= v_i-|N_i,t|^-1 _j _i,t v_j. The interaction-mesh objective is Emeshâ()=â„LtâVtâ()âLtâV~tâ„F2.E_mesh( q)= L_tV_t( q)-L_t V_t _F^2. (5) TERRA residuals. TERRA augments the interaction-mesh objective with segment- and terrain-interaction residual terms. The calibrated orientation residual iR=LogâĄ(Ri,tsrcâRiâ()â€) r_i^R=Log(R_i,t^srcR_i( q) ) matches source and target-model rotations at the pelvis and bilateral hip and knee sites. During stance, ixây=ixâyâ()âi r_i^xy= p_i^xy( q)- a_i holds each calcaneus at its touchdown anchor i a_i. Ankle and toe-displacement terms penalize all frame-to-frame horizontal motion and motion above 0.250.25 m/s. One-sided residuals [dminâdâĄ()]+[d_ -d( q)]_+ and [htââhsoleâ()]+[h_t -h_sole( q)]_+ penalize proximity between leftâright leg collision geometries and insufficient swing-sole clearance, respectively. Near terrain faces, vertical ankle and toe targets add at most 0.060.06 m of terrain-clearing lift. Calibrated sole offsets align source landmarks with the robotâs contact surface. During detected seated rests, a signed-distance residual brings the posterior pelvis collision geometries to the reconstructed seat. Detailed hyperparameter values are included in the Supp. Video. Sequential quadratic program. At iteration n of frame t, we compute an update Îâââ89 q ^89 to the current configuration t(n) q_t^(n). The 89 generalized coordinates comprise three root translations, a unit quaternion, and 82 joint coordinates. Let âŹt(n)=[lbât(n),ubât(n)]B_t^(n)=[ q^lb- q_t^(n), q^ub- q_t^(n)]: Îât(n)=argâminÎâ q_t^(n)= *arg\,min_ q wLââL+JLâÎââ22+âÎââtâW2 w_L\| r_L+J_L q\|_2^2+\| q- d_t\|_W^2 (6) +ât(n)+ÎââQ2 +\| q_t^(n)+ q\|_Q^2 +âmââłtwmâ„JmÎâm,tâ„22 + _m _tw_m\|J_m q- e_m,t\|_2^2 s.t. .t. ÏÂŻa+JÏ,aÎâ„âÏ”pen,aât, Ï_a+J_Ï,a qâ„- _pen, a _t, ÎâââŹt(n)â©Î·â2. q _t^(n)â© _2. Here L=vecâĄ(LtâVtâ(t(n))âLtâV~t) r_L=vec\! (L_tV_t( q_t^(n))-L_t V_t ) is the current interaction-mesh residual and JL=âL/âJ_L=â r_L/â q is its Jacobian. Thus, L+JLâÎâ r_L+J_L q approximates the residual after the update. The vector t=tâ1âât(n) d_t= q_t-1 - q_t^(n) denotes the displacement from the current iteration to the previous-frame solution. The matrix W weights this temporal smoothing residual, and Q penalizes absolute trunk angles. The set âłtM_t constitutes the active TERRA residuals described above, and tC_t contains active robotâenvironment geometry pairs. For each pair a, Ïa _a is its signed separation distance, with Ïa<0 _a<0 denoting penetration. We use the clipped distance ÏÂŻa=maxâĄÏa,â(Ï”pen+Ï) Ï_a= \ _a,-( _pen+Ï)\ with Ï”pen=0.9 _pen=0.9 m and per-iteration recovery cap Ï=10Ï=10 m. Thus, an existing deep penetration requests at most 10 m of correction in one SQP iteration rather than making the subproblem arbitrarily large. After each SQP solve, the root quaternion is radially projected onto the unit 3-sphere. Native Clarabel is used, with the condensed CVXPY formulation as a numerical fallback. To warm-start the solution, we append 30 copies of the first target frame and discard them afterward. The first solver frame uses trust radius 1.01.0 and at most 50 iterations; subsequent frames use radius 0.20.2 and at most 10 iterations. Finally, short collision outliers and tendon discontinuities are fixed via bounded interpolation after optimization. Fig. 3: Terrain-reconstruction. Method comparison on a Vielemeyer 10â10 descent ramp (top) and PRISM stepping boxes (bottom). Black crosses denote the same reference toe contacts in every panel, and dotted segments connect each contact to the reconstructed height at the same horizontal location. Vertical height is exaggerated for visibility. TABLE I: Terrain classification, reconstruction, and motionâterrain consistency across datasets. Entries are per-motion mean ± population standard deviation. (a) Known terrain All known terrain Gait120 Darmstadt Vielemeyer Method Family acc. (%) â (N=5,474N=5,474) Grade MAE (â) â (N=1190N=1190) Riser MAE (m) â (N=1184N=1184) Stool MAE (m) â (N=1102N=1102) Riser MAE (m) â (N=1282N=1282) Grade MAE (â) â (N=568N=568) Contact least squares N/A 1.683 ± 3.533 23.42 ± 8.71 490.00 ± 0.00 150.49 ± 57.41 1.199 ± 1.088 Voronoi N/A N/A 42.24 ± 39.54 80.90 ± 121.19 24.81 ± 36.39 N/A TERRA (without physical cues) 73.5 0.220 ± ± 0.235 12.25 ± 5.35 45.62 ± ± 13.57 7.12 ± 6.10 0.536 ± ± 0.333 (N=566) TERRA 99.9 0.223 ± 0.244 8.08 ± ± 6.99 45.62 ± ± 13.57 5.17 ± ± 4.36 0.536 ± ± 0.353 (b) PRISM mesh geometry Height MAE (m) â Footprint (%) â Method Success Foot (N=18N=18) Seat (N=13N=13) Observed (N=31N=31) Raised Flat IoU Contact least squares 31/31 102.41 ± 34.09 204.21 ± 19.14 145.10 ± 57.89 99.1 ± ± 5.0 37.1 ± 5.8 3.5 ± 1.0 Voronoi 31/31 34.13 ± 10.02 15.44 ± 8.38 26.29 ± 13.15 98.4 ± 4.6 40.3 ± 28.1 11.2 ± 17.9 TERRA 31/31 12.95 ± ± 6.56 11.16 ± ± 5.93 12.20 ± ± 6.36 90.8 ± 12.4 98.0 ± ± 0.6 47.3 ± ± 5.5 (c) AMASS-terrain motion consistency Method Success Contact MAE (m) â |re|â€50|r_e|†50 m (%) â Penetrating / floating (%) â Raised P / R / F1 (%) â Toeâankle pair MAE (m) â Free-space penetration (%) â Contact least squares 993/993 47.46 ± 30.38 57.5 34.5 / 8.0 46.1 / 99.3 / 62.9 24.73 ± 23.59 0.225 ± 0.856 Voronoi 966/993 25.57 ± 25.57 91.4 2.0 / 6.7 80.0 / 82.2 / 81.1 19.04 ± 21.31 0.000 ± ± 0.000 TERRA 993/993 10.24 ± ± 9.21 97.1 0.7 / 2.2 98.6 / 89.7 / 93.9 9.01 ± ± 7.46 0.077 ± 0.444 TABLE I: Retargeting performance across datasets and held-out policy-tracking performance. Retargeting Policy tracking Method Success DtenD_ten (%) â DcolD_col (%) â DpenD_pen (%) â DskatD_skat (%) â DfloatD_float (%) â Contact F1 (%) â Success (%) â MPJPE (m) â M-MoCap-Body 6478/6498 2.208 ± 3.852 12.09 ± 19.90 54.56 ± 35.53 4.86 ± 5.10 22.80 ± 36.78 92.86 ± 7.47 47.34 ± 0.22 90.26 ± 0.60 GMR 6498/6498 0.556 ± 1.087 21.82 ± 39.14 19.98 ± 24.19 9.25 ± 14.62 11.82 ± 11.27 74.19 ± 21.01 56.43 ± 0.64 91.97 ± 1.73 OmniRetarget 6392/6498 0.466 ± 1.243 4.98 ± 10.82 31.97 ± 29.22 12.70 ± 10.54 3.61 ± 9.79 86.05 ± 11.79 80.29 ± 1.39 87.23 ± 2.92 TERRA (no residuals) 6498/6498 0.134 ± 0.407 3.93 ± 8.45 33.45 ± 33.63 8.88 ± 7.03 4.18 ± 11.00 88.49 ± 9.51 86.52 ± 0.25 80.58 ± 2.31 TERRA+Voronoi 6470/6498 0.081 ± 0.285 0.54 ± 1.35 2.99 ± 14.97 2.63 ± 2.82 9.97 ± 18.73 92.96 ± 5.96 82.16 ± 0.41 71.65 ± ± 1.00 TERRA 6498/6498 0.026 ± ± 0.185 0.52 ± ± 1.34 0.16 ± ± 2.46 1.80 ± ± 1.87 1.02 ± ± 3.75 93.80 ± ± 4.06 87.59 ± ± 0.18 74.47 ± 1.51 I-D Policy Training We trained multi-motion, terrain-aware, muscle-actuated control policies on terrain-paired motions via motion imitation. The tracking policy observes joint positions and velocities, root height and projected gravity, heading-frame root velocity, muscle commands and activation states, four foot and toe touch sensors, and a heading-aligned Ă1111\!Ă\!11 height map centered on the pelvis at 0.10.1 m spacing. The one-step target encodes heading-frame root and mimic-site errors in position, orientation, and velocity; site positions and velocities are relative to the pelvis. Future targets at 0.2, 0.4, 0.6, and 0.8 s encode root motion and pelvis-relative site positions without motion phase. Training was performed with PPO on MJXâWarp [39]. The actor and critic are layer-normalized gated-residual MLPs. We used 8192 parallel environments at a control frequency of 100 Hz with five 2 ms physics steps per action. We used adaptive motion sampling to increase the sampling likelihood of hard motions. The PPO reward combines full-body pose and velocity tracking with a pelvis-relative upper-body position term and a terminal quality bonus. Small penalties discourage out-of-bounds and rapidly changing actions. In addition, a muscle activity regularization term discourages excessive activation while preventing muscle-unit silencing. Episodes terminate when the global or core-upper-body mean site error exceeds 0.150.15 m or the root-orientation error exceeds 11 rad. Each policy was trained for 4 billion steps. Hyperparameters are reported in Table IV. TABLE IV: Key numerical parameters for reconstruction, retargeting, and policy training. Parameter Value Stationary contacts vs=0.30v_s=0.30 m/s; nmin=5n_ =5; We=±0.30W_e=± 0.30 s; Ï”loc=0.05 _ loc=0.05 m. Geometry margins Contact expansion 0.10/0.050.10/0.05 m; maximum extension 0.00/0.150.00/0.15 m; free-space clearance 0.03/0.020.03/0.02 m (vertical/horizontal). TERRA weights Orientation 0.50.5; anchor 100100; velocity/displacement 150/200150/200; height 400400; separation 20,00020,000; clearance/route 1,5001,500; seat 2,0002,000. Constraint thresholds Ï”pen=0.0009 _ pen=0.0009 m; recovery cap Ï=0.010Ï=0.010 m/iteration; leg separation 0.0020.002 m; collision repair 0.0050.005 m for â€3†3 frames. DtenD_ ten Fraction with Îâl/l0>maxâĄ(10âEMA,0.001) l/l_0> (10\,EMA,0.001) and >0.05>0.05. Actor/critic width 2048,512,512,512,512,512,5122048,512,512,512,512,512,512. PPO parameters 8,192 envs; 20 steps; 1 epoch; 128 minibatches; Îł=0.99Îł=0.99; λ=0.95λ=0.95; clip 0.2; action SD 0.3; LR Ă10â4âĂ10â58\!Ă\!10^-4â 8\!Ă\!10^-5. Reward weights Joint q/vq/v: 0.5/0.25; root p/R/vp/R/v: 1/0.5/0.5; rel. sites: 1/0.5/0.5; core 1; terminal 25; action rate 10â410^-4; activation/floor 2/1. Training 4B steps; ⌠20 hours on four A100 GPUs IV Experimental Protocol Datasets. We sought to evaluate the full TERRA pipeline on a broad set of reference motions, combining diverse terrain interaction and biomechanical relevance. To do so, we applied TERRA on five distinct motion capture datasets. We extracted 993 non-flat motions from AMASS (AMASS-terrain) for large-scale, diverse motion [10]. We additionally considered 31 PRISM [12] motions containing static terrain, including ground-truth object meshes serving as geometric references for terrain reconstruction evaluation. Finally, we included 3,476 Gait120 motions [14], 1,282 Darmstadt stair motions [11], and 716 Vielemeyer ramp motions [13] as openly available biomechanics datasets with known terrain geometry, enabling quantitative evaluation of terrain reconstruction accuracy and physiological comparison with human data. For policy experiments, we added 595 Gait120 level-walking clips and 975 AMASS flat motions inherited from Kinesis [6]. Motion selection is described in detail in the Supplementary Video. Fig. 4: A policy trained on TERRA motions can imitate held-out motion on various terrains, such as stairs, ramps, individual platforms, and chairs. Reconstruction comparisons. To evaluate TERRAâs terrain reconstruction performance, we compared it to two baselines: (i) contact least squares, which fits one affine plane to the four-point kinematic contacts; and (i) Voronoi, a height-field baseline adapted from TIP [20] and SceneBot [21]. We also ablated the effect of TERRAâs swing-clearance cues to highlight their effect on terrain prior selection. For a fair comparison on chair reconstruction, TERRA and Voronoi retained their own detection and reconstruction but shared the posterior-mesh seat-height rule from Sec. I-B. We evaluated all methods individually on every dataset, according to the available ground-truth information (Table I). For Gait120, Darmstadt, and Vielemeyer, terrain labels and dimensions are known; we therefore report terrain-family accuracy and MAE in ramp grade, stair-riser height, and stool height. For PRISM, reference meshes additionally enable foot, seat, and combined-support height MAE, raised- and flat-region coverage, and raised-footprint intersection over union (IoU). The family-accuracy denominator pools every labeled known-terrain clip: 3,4763,476 Gait120 +1,282+1,282 Darmstadt +716+716 Vielemeyer =5,474=5,474; the other columns are dataset-specific. Neither these labels nor apparatus dimensions enter reconstruction. The numerical constants encode declared landmark, temporal, or collision resolutions rather than nominal apparatus dimensions. We report confidence intervals using subject-clustered bootstrap samples and break down reconstruction performance separately across terrain conditions. We also assessed robustness to noise (100-ms correlated isotropic Gaussian landmark noise at 2, 5, or 10 m RMS), support interval removal (10%, 20%, and 30%), and threshold perturbations (80% and 120% of the contact-speed, level-clustering, free-space, and minimum-raised-height). Because AMASS lacks scene geometry, we evaluated motionâterrain consistency using contact and non-collision criteria [40]. For event e and probe jâĄ(e)j(e), re=hmâ(xe,ye)â(zeâÎŽjâĄ(e)),r_e=h_m(x_e,y_e)- (z_e- _j(e) ), (7) where ÎŽj _j is a source-only offset from the probeâs lowest contact cluster. We report contact-height MAE; consistency (|re|â€50|r_e|†50 m), penetrating, and floating event rates; raised-support precision/recall/F1 at 40 m; toeâankle relative-support MAE; and non-support penetration below 30 m. Continuous metrics are unweighted per-motion mean ± population SD. Fig. 5: Qualitative comparison of population-averaged physiological profiles on held-out motions. Every human and policy trace was independently minâmax normalized, yielding a waveform-shape comparison. Shading denotes between-subject SEM. VL: vastus lateralis; VM: vastus medialis; SOL: soleus; GAS: gastrocnemius; BF: biceps femoris; TA: tibialis anterior. Retargeting comparisons. We compared TERRA with OmniRetarget [27], GMR [24], and M-MoCap-Body [7]. Since these methods do not support terrain reconstruction, we used the same TERRA scene across all baselines (Table I). We used a 100 Hz retargeting framerate across all methods. For OmniRetarget, we replaced the robot model with MyoFullBody, rebuilt its bounds for all 82 scalar joint coordinates, and provided the fitted SMPL-H shape, 17 landmarkâbody correspondences, and calibrated local offsets, retaining the original interaction-mesh, object-nonpenetration, and foot-sticking terms. For GMR, we followed the implementation from MuscleMimic [7], providing the actual terrain height for z-calibration. M-MoCap-Body used MuscleMimicâs native MyoFullBody fitter. We furthermore ablated TERRAâs residual terms to isolate their effect from the solver backend and other processing steps. Finally, to highlight the effect of terrain reconstruction on downstream motion retargeting, we added a TERRA+Voronoi baseline which combines Voronoi terrain reconstruction with TERRAâs motion retargeting pipeline. We define reference quality by preservation of task-relevant contacts together with satisfaction of target-specific anatomical and environmental constraints. We evaluate source-contact timing, tendon continuity, self-collision, penetration, skating, floating, and retargeting coverage. Coverage denotes converged retargeting runs over all motion inputs. We express each violation metric as a percentage of the relevant motion interval: the full motion interval for general constraints and source-derived contact or stance intervals for foot-specific constraints. We report DtenD_ ten when an adaptive tendon-length event exceeds 0.05 relative rest length; skating when the corresponding robot ankle or toe moves horizontally faster than 0.300.30 m/s, floating when a stance foot is more than 2020 m above its assigned surface, inter-leg collision when penetration exceeds 11 m, and environment penetration when penetration exceeds 1010 m. Contact preservation is the duration-weighted F1 score between source and retargeted kinematic-contact labels over the left and right ankles and toes. We apply the same contact detection criteria to both motions: probe speed below 0.300.30 m/s, height within 5050 m of the local minimum in a ±0.30± 0.30 s window, and a minimum contact duration of 0.150.15 s. True positives, false positives, and false negatives are accumulated over frameâprobe duration, and F1=2âTP/(2âTP+FP+FN)F_1=2TP/(2TP+FP+FN). Source labels are sampled at each methodâs declared output timestamps, with no temporal shift or matching tolerance. Dataset results are pooled by motion count and reported as the unweighted per-motion mean ± population SD. Tracking policy comparisons. To assess downstream executability, we compared TERRA to its retargeting baselines by training a PPO policy from each methodâs retargeted references, using the common setup (Sec. I-D). We partitioned the 8,068 source clips deterministically by recorded identity (subject, or AMASS actor/session), targeting a 15% test fraction while balancing dataset and motion type. For each method, we trained three independent policies with random seeds. The comparison was performed on the held-out test motions successfully retargeted by all methods. We evaluated each motion clip with five stochastic runs. Success denotes trajectory completion within a tracking termination bound of 0.250.25m, and tracking error is average global mimic-site MPJPE over executed control steps. Physiological evaluation. We used held-out recordings from Gait120 [14], Darmstadt [11], and Vielemeyer [13] to visualize policy-generated EMG and vertical-GRF waveforms alongside human data. Each case used ten stochastic rollouts and a 0.250.25-m termination threshold; every completed gait was retained, including gaits from episodes that later terminated. Each completed policy gait and each measured subject trace were independently minâmax normalized to compare waveform shape, focusing the analysis on temporal profiles. We averaged policy gaits within subject and then averaged policy and measured traces separately across subjects in the same condition. We selected level walking, ramp descent, stair descent, and sit/stand to span four task families with available EMG and GRF profiles and, for each, show vertical GRF and three representative muscles (Fig. 5). V Results V-A Terrain Reconstruction Qualitatively, TERRAâs priors enabled the method to flexibly infer slanted or piecewise-flat surfaces, whereas baselines methods were confined to more rigid representations (Fig. 3). We quantitatively evaluated the terrain reconstruction quality of TERRA by asking two key questions: how well does TERRA discriminate between different terrain types, and how closely does the inferred terrain geometry match the real geometry? Overall, TERRA achieved 99.9% terrain-family accuracy, successfully classifying ramps from stairs across datasets and conditions. Importantly, this classification accuracy was paired with strong reconstruction fidelity, namely sub-centimeter stair height accuracy, and sub-degree ramp accuracy, while outperforming baselines on all terrain categories (Table I). Removing the foot-orientation and swing-clearance cues reduced family accuracy to 73.5%, while having little effect on other metrics (performance is identical to TERRA for Tables IIb,c and thus not shown). The subject-clustered 95% CI for pooled family accuracy was 99.853â99.982% (144 datasetâsubject clusters). Across independent captures, grade MAE was 0.544â/0.529â at 7.5â/10â, and riser MAE was 7.36/4.34/3.61 m at 100/170/240 m. TERRA remained robust to threshold perturbations and landmark noise, while substantial support removal caused a gradual, terrain-dependent degradation, most notably for stools (Table V). TABLE V: Terrain reconstruction accuracy (%) across perturbations. Level Ramp Stairs Stool All (148) (1,758) (2,466) (1,102) (5,474) Clean 100.00 100.00 99.84 100.00 99.93 Noise 2 m 100.00 99.98 99.81 100.00 99.91 Noise 5 m 100.00 99.75 99.10 100.00 99.51 Noise 10 m 99.59 95.78 96.52 99.96 97.06 Support â10%-10\% 100.00 98.76 99.61 89.46 97.30 Support â20%-20\% 100.00 96.64 98.96 79.84 94.40 Support â30%-30\% 100.00 91.18 97.42 70.36 90.04 Thresholds 80%80\% 100.00 99.89 99.96 100.00 99.95 Thresholds 120%120\% 100.00 100.00 99.80 100.00 99.91 V-B Motion Retargeting We tested TERRA and its baselines on source-contact preservation as well as anatomical and terrain-interaction validity. TERRA achieved the best results on all five physical violation metrics and contact-timing F1 (Table I). TERRAâs residual optimization terms raised contact-timing F1 from 88.49% to 93.80% while lowering all five violation metricsâespecially terrain penetration, which dropped from 33.45% to 0.16%. Compared with TERRA+Voronoi, the reconstructed TERRA terrain further reduced penetration, skating, and floating while increasing Contact F1. V-C Tracking Policy Performance Qualitatively the policy performed well across different affordances (Fig. 4). We next asked whether improvements in reference validity were accompanied by improved downstream imitation performance. In the held-out evaluation setting, TERRA-retargeted reference motions produced the highest completion rate compared to baselines (Table I); TERRA+Voronoi achieved better MPJPE but with a lower success rate. Broken down by terrain family, TERRA achieved the highest success rate on flat ground, ramps, boxes, and seats, and the lowest MPJPE on flat ground, ramps, and boxes (Table VI), while TERRA+Voronoi performed best on stairs and attained the lowest seat MPJPE. We encourage readers to view the supplementary video for in-depth qualitative results. TABLE VI: Held-out policy tracking by terrain family. Terrain (N) OmniRetarget TERRA+Voronoi TERRA Success rate (%) Flat (262) 88.07 ± 0.62 91.09 ± 0.75 91.70 ± ± 0.46 Ramp (286) 98.09 ± 0.08 89.18 ± 1.06 99.49 ± ± 0.18 Stairs (422) 74.14 ± 3.49 85.01 ± ± 0.24 81.90 ± 1.73 Boxes (61) 38.91 ± 4.64 39.45 ± 0.19 46.45 ± ± 4.45 Seats (246) 72.11 ± 0.61 70.19 ± 0.38 89.32 ± ± 0.82 MPJPE (m) Flat (262) 82.42 ± 3.13 69.97 ± ± 1.57 70.01 ± 2.55 Ramp (286) 75.82 ± 3.36 67.83 ± 1.02 66.74 ± ± 2.92 Stairs (422) 85.18 ± 3.06 66.65 ± ± 1.12 69.50 ± 2.57 Boxes (61) 94.02 ± 2.26 85.25 ± 0.55 82.50 ± ± 0.77 Seats (246) 107.45 ± 2.48 83.09 ± ± 1.13 94.75 ± 2.50 V-D Physiological Evaluation Terrain-paired biomechanics datasets let us compare signals produced by learned policies directly with measured human profiles for the same locomotion condition. Across the selected held-out flat, ramp, stair, and sit/stand examples, the policy and human profiles show both similarities and visible mismatches in EMG timing and shape (Fig. 5). The vertical-GRF panels provide a visual comparison of stance timing and normalized waveform shape. Taken together, these examples suggest that TERRA can serve as a tool to study physiological processes in the future. VI Conclusion TERRA demonstrates that motion kinematics can provide enough environmental evidence to recover task-relevant collision geometry for muscle-actuated control. Across stairs, ramps, isolated supports, and chairs, TERRA reduced anatomical and interaction violations and yielded the highest held-out completion rate. By incorporating diverse terrain navigation into the behavioral repertoire of musculoskeletal control policies, TERRA may open a novel route to studying affordance representation in embodied agents. For instance, classical findings that stair climbing and sitting are perceived in body-scaled terms [41, 42] could be revisited by relating learned terrain representations to the musculoskeletal capabilities of the body, complementing neural accounts of how the brain specifies competing action opportunities [43, 2]. Dynamic primitive geometry, online retargeting, and moving surfaces remain as open challenges for future work. References [1] J. J. Gibson, The Ecological Approach to Visual Perception. Boston, MA: Houghton Mifflin, 1979. [2] D. E. Angelaki, A. Batista, T. Fitzgerald, J. F. Kominsky, M. Lengyel, A. Mathis, M. W. Mathis, C. F. Moss, C. M. Niell, J.-P. Noel, et al., âThe simons collaboration on ecological neuroscience: Studying how the brain interacts with the world,â Neuron, 2026. [3] J. M. Wang, S. R. Hamner, S. L. Delp, and V. Koltun, âOptimizing locomotion controllers using biologically-based actuators and objectives,â ACM Transactions on Graphics (TOG), vol. 31, no. 4, p. 1â11, 2012. [4] Y. Lee, M. S. Park, T. Kwon, and J. Lee, âLocomotion control for many-muscle humanoids,â ACM Transactions on Graphics (TOG), vol. 33, no. 6, p. 1â11, 2014. [5] Y. Feng, X. Xu, and L. Liu, âMusclevae: Model-based controllers of muscle-actuated characters,â in SIGGRAPH Asia 2023 Conference Papers, 2023, p. 1â11. [6] M. Simos, A. S. Chiappa, and A. Mathis, âKinesis: Motion imitation for human musculoskeletal locomotion,â ICRA, 2026. [7] C. Li, C. Wang, B. Ziliotto, M. Simos, J. Kovecses, G. Durandau, and A. Mathis, âTowards embodied AI with MuscleMimic: Unlocking full-body musculoskeletal motor learning at scale,â arXiv preprint arXiv:2603.25544, 2026. [8] C. Wang, C. K. Tan, B. K. Hodossy, E. Lyu, J. Guo, W. Zhao, H. Liu, C. Li, M. Simos, B. Ziliotto, et al., âMyochallenge 2025: A new benchmark for human athletic intelligence,â arXiv preprint arXiv:2605.15650, 2026. [9] Y. Wei, C. Zuo, S. Zhuang, H. Gong, Y. Liu, and Y. Sui, âScaling whole-body human musculoskeletal behavior emulation for specificity and diversity,â arXiv preprint arXiv:2603.29332, 2026. [10] N. Mahmood, N. Ghorbani, N. F. Troje, G. Pons-Moll, and M. J. Black, âAmass: Archive of motion capture as surface shapes,â in Proceedings of the IEEE/CVF international conference on computer vision, 2019, p. 5442â5451. [11] M. Grimmer, G. Zhao, J. Zeiss, F. Weigand, S. Lamm, M. Steil, and A. Heller, âDarmstadt stair ambulation dataset including level walking, stair ascent, stair descent and gait transitions at three stair heights,â TUdatalib dataset, 2023. [12] R. Hori, J.-T. Song, Z. Luo, J. Cao, S. Shin, H. Saito, and K. Kitani, âGround reaction inertial poser: Physics-based human motion capture from sparse IMUs and insole pressure sensors,â in Proceedings of the IEEE/CVF Conference on Computer Vision and Pattern Recognition (CVPR), June 2026, p. 28 435â28 445. [13] J. Vielemeyer, L. Tronicke, L. Schreff, R. Abel, K. Lechler, and R. MĂŒller, âA full-body motion capture gait dataset of healthy young adults walking ramps up and down,â Scientific Data, vol. 13, p. 18, 2026. [14] J. Boo, D. Seo, M. Kim, and S. Koo, âComprehensive human locomotion and electromyography dataset: Gait120,â Scientific Data, vol. 12, p. 1023, 2025. [15] H. Yi, C.-H. P. Huang, S. Tripathi, L. Hering, J. Thies, and M. J. Black, âMime: Human-aware 3d scene generation,â in 2023 IEEE/CVF Conference on Computer Vision and Pattern Recognition (CVPR). IEEE, 2023, p. 12 965â12 976. [16] J. Li, T. Huang, Q. Zhu, and T.-T. Wong, âPhysics-based scene layout generation from human motion,â in ACM SIGGRAPH 2024 Conference Papers, 2024, p. 1â10. [17] A. Allshire, H. Choi, J. Zhang, D. McAllister, A. Zhang, C. M. Kim, T. Darrell, P. Abbeel, J. Malik, and A. Kanazawa, âVisual imitation enables contextual humanoid control,â arXiv preprint arXiv:2505.03729, 2025. [18] Z. Wang, J. Wang, J. Tan, Y. Zhao, J. Hodgins, S. Tulsiani, and D. Ramanan, âContact-guided real2sim from monocular video with planar scene primitives,â in International Conference on Learning Representations, vol. 2026, 2026, p. 115 452â115 468. [19] Q. Zhang, J. Ma, P. Liu, S. Shi, Z. Su, Z. Wang, J. Sun, W. Cui, J. Yu, G. Han, et al., âMeshmimic: Geometry-aware humanoid motion learning through 3d scene reconstruction,â arXiv preprint arXiv:2602.15733, 2026. [20] Y. Jiang, Y. Ye, D. Gopinath, J. Won, A. W. Winkler, and C. K. Liu, âTransformer inertial poser: Real-time human motion reconstruction from sparse imus with simultaneous terrain generation,â in SIGGRAPH Asia 2022 Conference Papers, 2022, p. 1â9. [21] S. Chen, S. Zhao, Z. Wu, J. Li, G. Shi, and C. K. Liu, âSceneBot: Contact-prompted general humanoid whole body tracking with scene-interaction,â arXiv preprint arXiv:2606.27581, 2026. [22] Bones Studio, âBONES-SEED: Skeletal everyday embodiment dataset,â https://huggingface.co/datasets/bones-studio/seed, 2025. [23] K. Lee, S. Kim, M. Park, H. Kim, D. Hwang, H. Lee, and J. Choo, âPhuma: Physically-grounded humanoid locomotion dataset,â arXiv e-prints, p. arXivâ2510, 2025. [24] J. P. Araujo, Y. Ze, P. Xu, J. Wu, and C. K. Liu, âRetargeting matters: General motion retargeting for humanoid motion tracking,â arXiv preprint arXiv:2510.02252, 2025. [25] T. Huang, F. Yuan, J. Gu, S. Fang, X. Zhang, Y. Wang, W. Gao, and S. Zhang, âHuman2humanoid: Physics-aware cross-morphology motion retargeting for humanoid robots,â arXiv preprint arXiv:2606.03476, 2026. [26] H. Wang, Q. Liao, B. Zhang, K. Ren, K. Sreenath, and X. Xiong, âSpark: Skeleton-parameter aligned retargeting on humanoid robots with kinodynamic trajectory optimization,â arXiv preprint arXiv:2603.11480, 2026. [27] L. Yang, X. Huang, Z. Wu, A. Kanazawa, P. Abbeel, C. Sferrazza, C. K. Liu, R. Duan, and G. Shi, âOmniretarget: Interaction-preserving data generation for humanoid whole-body loco-manipulation and scene interaction,â arXiv preprint arXiv:2509.26633, 2025. [28] S. Lee, M. Park, K. Lee, and J. Lee, âScalable muscle-actuated human simulation and control,â ACM Transactions On Graphics (TOG), vol. 38, no. 4, p. 1â13, 2019. [29] A. Seth, M. Sherman, J. A. Reinbolt, and S. L. Delp, âOpenSim: A musculoskeletal modeling and simulation framework for in silico investigations and exchange,â Procedia Iutam, vol. 2, p. 212â232, 2011. [30] T. Geijtenbeek, âScone: Open source software for predictive simulation of biological motion,â Journal of Open Source Software, vol. 4, no. 38, p. 1421, 2019. [31] V. Caggiano, H. Wang, G. Durandau, M. Sartori, and V. Kumar, âMyosuiteâa contact-rich simulation suite for musculoskeletal motor control,â arXiv preprint arXiv:2205.13600, 2022. [32] A. S. Chiappa, A. Marin Vargas, A. Huang, and A. Mathis, âLatent exploration for reinforcement learning,â Advances in Neural Information Processing Systems, vol. 36, p. 56 508â56 530, 2023. [33] K. He, C. Zuo, C. Ma, and Y. Sui, âDynsyn: Dynamical synergistic representation for efficient learning and control in overactuated embodied systems,â arXiv preprint arXiv:2407.11472, 2024. [34] Y. Wei, C. Zuo, and Y. Sui, âScalable exploration for high-dimensional continuous control via value-guided flow,â arXiv preprint arXiv:2601.19707, 2026. [35] P. Schumacher, T. Geijtenbeek, V. Caggiano, V. Kumar, S. Schmitt, G. Martius, and D. F. Haeufle, âEmergence of natural and robust bipedal walking by learning from biologically plausible objectives,â iScience, 2025. [36] B. An, A. S. Chiappa, M. Simos, C. Li, and A. Mathis, âArnold: A multi-task, multi-embodiment muscle transformer policy,â 2026. [37] W. Wang, N. Bianco, G. Tevet, J. Hicks, C. K. Liu, S. Delp, and K. Fatahalian, âLearning realistic athletic sprinting without demonstrations,â in SIGGRAPH Asia 2026 Conference Papers, 2026. [38] J. Romero, D. Tzionas, and M. J. Black, âEmbodied hands: Modeling and capturing hands and bodies together,â ACM Transactions on Graphics, vol. 36, no. 6, p. 245:1â245:17, Nov. 2017. [39] J. Schulman, F. Wolski, P. Dhariwal, A. Radford, and O. Klimov, âProximal policy optimization algorithms,â arXiv preprint arXiv:1707.06347, 2017. [40] M. Hassan, P. Ghosh, J. Tesch, D. Tzionas, and M. J. Black, âPopulating 3d scenes by learning humanâscene interaction,â in Proceedings of the IEEE/CVF Conference on Computer Vision and Pattern Recognition (CVPR). IEEE, June 2021, p. 14 708â14 718. [41] W. H. Warren, âPerceiving affordances: visual guidance of stair climbing.â Journal of experimental psychology: Human perception and performance, vol. 10, no. 5, p. 683, 1984. [42] L. S. Mark, âEyeheight-scaled information about affordances: a study of sitting and stair climbing.â Journal of experimental psychology: human perception and performance, vol. 13, no. 3, p. 361, 1987. [43] P. Cisek and J. F. Kalaska, âNeural mechanisms for interacting with a world full of action choices,â Annual review of neuroscience, vol. 33, no. 1, p. 269â298, 2010.